Gray Leaf Spot

Gray leaf spot is a destructive disease that primarily affects Stenotaphrum secundatum but has also been observed in other turfgrass species. While it has been reported on Cynodon and Paspalum spp., they are not considered primary hosts in the field. The disease is commonly known as gray leaf spot in North America and other regions, and it is also referred to as “blast” disease in forage grasses. Blast disease, caused by a closely related pathogen, affects rice, wheat, and finger millet among other cereal grains.

 

Credit: PACE Turf

 

Causal agents

Pyricularia oryzae fungus

 

Primary Hosts

Originally a disease of Stenotaphrum secundatum (St. Augustinegrass). It has become very destructive to Lolium perenne (perennial ryegrass) and Festuca arundinaceae (tall fescue) as well. 

 

Disease Cycle

The gray leaf spot pathogen, Pyricularia oryzae, survives unfavorable conditions as dormant mycelium in infected leaves and plant debris. When the environment becomes suitable for its growth, the fungus produces conidiophores on necrotic tissue. These conidiophores generate and release conidia, which can be dispersed by wind, water, machinery, and animals. Conidia are produced abundantly in warm and humid conditions. Germination and infection of host tissue occur when the relative humidity is high and the leaf surface is moist. However, excessive leaf wetness can inhibit spore production. Ideal conditions for spore production and subsequent infection involve alternating wet and dry cycles over several days.

 

Credit: Wakar Uddin/Penn State University; Accessible Version (PDF)

Figure 14.7.1: Diagram of the Gray leaf spot disease cycle.
 
 

Epidemiology

Gray leaf spot is most severe in warm and humid weather, with optimal temperatures ranging from 28 to 32°C (82 to 90°F). It commonly affects Lolium spp. during the summer months in the transition zone through the mid-Atlantic region. Outbreaks can last until October in the eastern United States and are active from late spring to early summer in the southern United States during overseeding. Newly established plantings are more susceptible, especially with moderate to high nitrogen fertility.

Factors contributing to disease development include:

  • mowing height
  • temperature
  • leaf wetness
  • herbicides
  • drought

A predictive model based on temperature and leaf wetness is available for management.

 

Symptoms

Symptoms in Lolium spp. and Festuca arundinacea include small, water-soaked lesions that turn necrotic, varying in color and shape. Lesions may coalesce, blighting the leaf blades and forming irregular shapes. Infected patches can rapidly expand, resembling symptoms of brown patch. In severe cases, the entire turf may die. Festuca arundinacea generally recovers more rapidly than Lolium spp. when conditions are no longer favorable for disease development.

Symptoms of gray leaf spot in warm-season turfgrasses include dark, water-soaked lesions on leaves and stems that enlarge into round to diamond-shaped spots. The spots often occur along the leaf edges or midrib and may extend across the entire leaf. They have tan to gray centers with distinct reddish-brown borders.

 

Credit: Peter Dernoeden/University of Maryland

Figure 14.7.2: Gray leaf spot symptoms vary, but can appear as (a) a rapid decline of entire swards of perennial ryegrass or (b) as a general brown or thinned-out area on tall fescue. Lesions are also variable and often are confused with Bipolaris diseases. Lesions may appear as (c) brown or black, but more often (d) have a yellow halo surrounding a tan central portion with a brown border. Because symptoms can be confused with other diseases, positive identification of the conidia is required for an accurate diagnosis.

 

 

Signs

In both cool- and warm-season turfgrasses during warm and humid weather, the spots may be covered in grayish, felty masses of conidia. Individual conidia are borne on gray-to-olive conidiophores and are hyaline, pyriform in shape with two septations. Although perithecia have been produced in mating studies, the sexual stage of the pathogen has not been observed in nature.

 

Credit: Peter Dernoeden/University of Maryland and John Kaminski

Figure 14.7.3: (a&b) Signs of the pathogen include mycelium, but positive diagnosis can be made by observing a fuzzy gray spore mass around lesions. Conidia are borne on (c) conidiophores and (d) individual conidia have 2 septations and are tear-drop shaped.
 
 

Management

Managing gray leaf spot in turfgrass requires a combination of cultural and chemical approaches. To minimize the impact of the disease, it is important to implement cultural practices that reduce stress on the turf. This includes avoiding prolonged leaf wetness, excessive nitrogen application, soil compaction, and stress caused by plant growth regulators or herbicides. Deep and infrequent watering is recommended to prevent drought stress. Delaying overseeding in cooler weather can help reduce disease severity. While the impact of clippings removal is limited, it may be beneficial in low disease severity situations. Preventive fungicide applications are recommended in situations where the disease is chronic.

 

Cultural

To manage gray leaf spot, it is important to implement practices that minimize stress on the turf. This includes avoiding the following:

  • extended periods of leaf wetness
  • excessive nitrogen application
  • soil compaction
  • stress induced by plant growth regulators or herbicides
During hot, humid weather, limit the use of quick-release, water-soluble nitrogen fertilizers. Water the turf deeply and infrequently to prevent drought stress.

Delay overseeding in cooler autumn weather to reduce disease severity. While clippings removal has limited impact, it may be beneficial if disease severity is low.

 

Chemical

Fungicides can be effective in controlling gray leaf spot, especially for juvenile turfgrass during sod growth, or problematic areas. Preventive fungicide applications are recommended in chronic disease situations. Be aware of fungicide resistance and follow appropriate resistance management strategies, as some isolates of P. oryzae have shown resistance to fungicides within the QoI fungicide class. Where resistance isn’t an issue, the QoI fungicides work well. Additional fungicides include DMIs, thiophanate-methyl and combinations of these fungicides with chlorothalonil.